Publications

Grether, G.F, and Hopkins, E. L. Grether, G.F. 2025. Sexual selection and speciation. In: Encyclopedia of Animal Behavior,  (3rd ed.). Ed. R. Glog, Elsevier, Academic Press, in press pdf

Hoffman, S., Benson, L., Philson, C.S., Chock, R.Y., Curti, J.N., Flores-Negrón, C.F. and Grether, G.F. 2024 Reduction in mutualistic ant aggressive behavior upon sugar supplementation Biotropica e13390. DOI: 10.1111/btp.13390 pdf

Grether, G.F., Finneran, A.E., and Drury, J.P. 2024 Niche differentiation, reproductive interference, and range expansion. Ecology Letters DOI: 10.1111/ele.14350 pdf

McEachin, S. Drury, J.P., and Grether G.F. 2024 Competitive displacement and agonistic character displacement, or the ghost of interference competition. American Naturalist DOI: 10.1086/728671 pdf

Snowden, S.K., Shier, D.M., Stemp, K.M., and Grether, G.F. 2024 Assessing the potential for successful translocation and co-management of two endangered aquatic species. Animal Conservation DOI:10.1111/acv.12893 pdf

Patterson, C.W., Bonillas-Monge, E., Brennan, A., Grether, G.F., Mendoza-Cuenca, L., Tucker, R., Vega-Sánchez, Y. Drury, J. 2023 A chromosome-level genome assembly for the smoky rubyspot damselfly (Hetaerina titia). Journal of Heredity DOI: 10.1093/jhered/esad070 pdf

Keith, S.A., Drury, J.P., McGill, B.J., and Grether, G.F. 2023 Macrobehaviour: behavioural variation across space, time and taxa. Trends in Ecology & Evolution DOI: 10.1016/j.tree.2023.08.007 pdf

Shier, D.M., Grether, G.F., Reid-Wainscoat, E.E., Fox, S., Pemberton, D., Elmer, J.  and Swaisgood, R.R. 2023 Investigating scent communication and latrine use to inform translocation of the Tasmanian devil. Behavioral Ecology & Sociobiology DOI: 10.1007/s00265-023-03371-1 pdf

Grether, G.F., Beninde, J., Beraut, E., Chumchim, N., Escalona, MacDonald, Z.G., Miller, C., M., Sahasrabudhe, R., Shedlock, A.M., Toffelmier, E., Shaffer, H.B. 2023 Reference genome for the American rubyspot damselfly, Hetaerina americana.  Journal of Heredity DOI: 10.1093/jhered/esad031 pdf

Nesbit, D.A., Cowen, M.C., Grether, G.F., and Drury, J.P. 2023 Interspecific territoriality has facilitated recent increases in the breeding habitat overlap of North American passerines. Ecography e06573 DOI: 10.1111/ecog.06573 pdf

Standring, S., Sánchez-Herrera, M., Guillermo-Ferreira, R., Ware, J.L., Vega-Sánchez, Y.M., Clement, R., Drury, J.P., Grether, G.F., González-Rodríguez, Mendoza-Cuenca, L., Bota-Sierra, C.A., Bybee, S. 2022 Evolution and biogeographic history of rubyspot damselflies (Hetaerininae: Calopterygidae: Odonata). Diversity 14:757 DOI: 10.3390/d14090757 pdf

Grether, G.F. and Okamoto, K.W. 2022 Eco-evolutionary dynamics of interference competition. Ecology Letters DOI: 10.1111/ele.14091 pdf

Wu, C., Bhagawat, C., Goldman, M.R., Punjabi, N.A., Shier, D.M., Arauco-Aliaga, R.P., and Grether, G.F. 2022 Aggression in a Western Amazonian colonial spider, Philoponella republicana. Journal of Arachnology 50: 277-287 DOI: 10.1636/JoA-S-20-093 pdf

Grether, G.F., Siepielski, A.M., and Gómez-Llano, M. 2022 Ecological differentiation, interference, and coexistence in odonates. In: Dragonflies and damselflies: Model organisms for ecological and evolutionary research, Second Edition. Eds. A. Córdoba-Aguilar, C.D. Beatty, and J.T. Bried, Oxford University Press. DOI: 10.1093/oso/9780192898623.003.0015 pdf

Chock, R.Y., Shier, D.M., and Grether, G.F. 2022 Niche partitioning in an assemblage of granivorous rodents, and the challenge of community-level conservation. Oecologia  DOI: 10.1007/s00442-021-05104-5 pdf

McEachin, S., Drury, J.P., Anderson, C.N., Grether, G.F. 2021 Mechanisms of reduced interspecific interference between territorial species. Behavioral Ecology DOI: 10.1093/beheco/arab115 pdf

Grether, G.F. 2021 Exciting discoveries in animal signaling and sexual dichromatism. Functional Ecology 35:1624–1626. pdf

Gonzalez-Karlsson, A. and Grether, G.F. 2021 Trade-off mediated by pyrrolizidine alkaloids predicts alternative reproductive tactics in Ithomiine butterflies. Ecological Entomology DOI: 10.1111/een.13055 pdf

Grether, G.F. 2021 Developing and testing curricula for teaching evolutionary concepts at the elementary school level. American Biology Teacher 83: 96-103. pdf

Grether, G.F., Chock, R.Y., Cowen, M.C., De La Cruz-Sevilla, J.S., Drake, T.N., Lum, K.S., Ovakimyan, A., Sood, S.R., Tang, W., Whitney, K.A. and Yang, M.Z. 2021 Teaching evolutionary concepts in elementary school. American Biology Teacher 83: 118-119. pdf

Cowen, M.C., Drury, J.P. and Grether, G.F. 2020 Multiple routes to interspecific territoriality in sister species of North American perching birds. Evolution 74-9: 2134–2148 DOI:10.1111/evo.14068 pdf

Drury, J.P., Cowen, M.C. and Grether, G.F. 2020 Competition and hybridization drive interspecific territoriality in birds. Proceedings of the National Academy of Sciences DOI: 10.1073/pnas.1921380117 pdf

Grether  G.F., Drury, J.P., Okamoto, K.W., McEachin, S. and Anderson, C.N. 2020. Predicting evolutionary responses to interspecific interference in the wild. Ecology Letters 23: 221–230 DOI: 10.1111/ele.13395 pdf

Kirschel, A.N.G., Nwankwo, E.C. Seal, N., and Grether, G.F. 2020. Time spent together and time spent apart affect song, feather colour and range overlap in tinkerbirds. Biological Journal of the Linnean Society 129:439-458. DOI: 10.1093/biolinnean/blz191 pdf

Grether, G.F. 2020. Convergent and divergent selection drive plumage evolution in woodpeckers. Nature Communications 11:124.  pdf

Chock, R.Y., Grether, G.F. and Shier, D.M 2019. Cache pilfering in a granivore guild: implications for reintroduction management. Journal of Wildlife Management 83: 1607-1616. DOI: 10.1002/jwmg.21736 pdf

Grether, G.F. 2019. Territoriality in aquatic insects.  In: Aquatic Insects – Behavior and Ecology, Eds. K. Del Claro and R. Guillermo. Springer. pdf

Drury, J.P., Anderson, C.N., Cabezas Castillo, M.B., Fisher, J., McEachin, S., and Grether, G.F. 2019. A general explanation for the persistence of reproductive interference. American Naturalist 194: 268-275 DOI: 10.1086/704102  pdf

Drury, J.P, Barnes, M., Finneran, A.E., Harris, M. and Grether, G.F. 2019. Continent-scale phenotype mapping using citizen scientists’ photographs. Ecography 42: 1436-1445. DOI: 10.1111/ecog.04469 pdf

Nwankwo, E.C., Mortega, K.G., Karageorgos, A., Ogolowa, B.O.,Grether, G. F.,Monadjem, A.,and Kirschel, A.N.G. 2019. Rampant introgressive hybridization in Pogoniulustinkerbirds despite millions of years of divergence. Biological Journal of the Linnean Society 127: 125–142. pdf

Grether, G.F.2019. Sexual Selection and Speciation. In: Choe, J.C. (Ed.),Encyclopedia of Animal Behavior (2nd ed.). vol. 4, pp. 591–598. Elsevier, Academic Press. pdf

Cabezas Castillo, M.B. & Grether, G.F.  2018. Why are female color polymorphisms rare in territorial damselflies? Ethology 124:667–673 DOI: 10.1111/eth.12771 pdf

Grether, G.F. Character Displacement.  2018. In: Oxford Bibliographies in Evolutionary Biology. Ed. K. Pfennig. New York: Oxford University Press. pdf

Chock, R.Y., Shier, D.M., & Grether, G.F2018. Body size, not phylogenetic relationship or residency, drives interspecific dominance in a little pocket mouse community. Animal Behaviour 137:197-204. pdf

Drury, J.P., Grether, G.F., Garland, Jr., T., & Morlon, H. 2017. An assessment of phylogenetic tools for analyzing the interplay between interspecific interactions and phenotypic evolution. Systematic Biology 67: 413–427. pdf

Grether G.F., Peiman K.S., Tobias J.A., Robinson B.W. 2017. Causes and consequences of behavioral interference between species. Trends Ecol. Evol. DOI: 10.1016/j.tree.2017.07.004 pdf

Harvey, B.D., Vanni, K.N., Shier, D.M., Grether, G.F. 2017. Experimental test of the mechanism underlying sexual segregation at communal roosts of harvestmen (Prionostemma spp.) Ethology 123:516–525. DOI: 10.1111/eth.12623 pdf

Grether, G.F. 2017. Sexual selection and speciation. In: Encyclopedia of Animal Behavior, M. Breed and J. Moore (eds.). vol., pp., Oxford: Academic Press.

Sandkam, B.A., Deere, K.A., Johnson, A.M., Grether, G.F., Rodd, F.H. and Fuller, R.C. 2016. Exploring visual plasticity: dietary carotenoids can change color vision in guppies (Poecilia reticulata). Journal of Comparative Physiology A. 202:527–534. DOI 10.1007/s00359-016-1097-9 pdf

Losin, N., Drury, J.P, Peiman, K.S., Storch, C. and Grether, G.F. 2016. The ecological and evolutionary stability of interspecific territoriality. Ecology Letters 19: 260–267. DOI: 10.1111/ele.12561 pdf

Drury, J.P., Anderson, C.N., and Grether, G.F. 2015. Seasonal polyphenism in wing coloration affects species recognition in rubyspot damselflies (Hetaerina spp.). Journal of Evolutionary Biology 28: 1439-1452. pdf

Grether, G.F., Drury, J.P., Berlin, E. and Anderson, C.N. 2015. The role of wing coloration in sex recognition and competitor recognition in rubyspot damselflies (Hetaerina spp.). Ethology 121: 674-685. pdf

Drury, J.P., Okamoto, K.W., Anderson, C.N., and Grether, G.F. 2015. Reproductive interference explains persistence of aggression between species. Proceedings of the Royal Society B. 282: 20142256. DOI: 10.1098/rspb.2014.2256 pdf

Ribeiro, R., McCormack, J., Alvarez, H., Carrasco, L., Grether, G.F., Mena, P., Sedano, R., Smith, T.B., and Karubian, J. 2015. Loss of sexual dimorphism is associated with loss of lekking behavior in the green manakin (Xenopipo holochora) Journal of Avian Biology 46: 307-314. DOI: 10.1111/jav.00545 pdf

Kemp, D.J. and Grether, G.F. 2015. Integrating functional and evolutionary approaches to the study of colour-based animal signals. In: Animal Signaling and Function: An Integrative Approach. D.J. Irschick, M. Briffa, and J. Podos (editors). John Wiley & Sons.

Drury, J.P and Grether, G.F 2014. Interspecific aggression, not interspecific mating, drives character displacement in the wing coloration of male rubyspot damselflies (Hetaerina). Proceedings of the Royal Society B. 281 20141737 DOI: 10.1098/rspb.2014.1737 pdf

Grether, G.F., Aller, T.L., Grucky, N.K., Antaky, C.C. Levi, A. and Townsend, V.R. Jr. 2014. Species differences and geographic variation in the communal roosting behavior of Prionostemma harvestmen in Central American rainforests.  Journal of Arachnology 42:257-267. pdf

Grether, G.F., Levi, A., Antaky, C. and Shier, D.M. 2014. Communal roosting sites are potential ecological traps: experimental evidence in a Neotropical harvestman. Behavioral Ecology and Sociobiology 68:1629–1638. DOI: 10.1007/s00265-014-1771-2 pdf

Grether, G.F. 2013. Redesigning the genetic architecture of phenotypically plastic traits in a changing environment.  Biological Journal of the Linnean Society 112: 276–286. DOI: 10.1111/bij.12064 pdf

Okamoto, K.W. and Grether, G.F. 2013. The evolution of species recognition in competitive and mating contexts: the relative efficacy of alternative mechanisms of character displacement. Ecology Letters 16: 670–678. DOI: 10.1111/ele.12100 pdf

Grether, G.F., Anderson, C.N., Drury, J.P., Kirschel, A.N.G., Losin, N., Okamoto, K. and Peiman, K. 2013. The evolutionary consequences of interspecific aggression. Annals of the New York Academy of Sciences, The Year in Evolutionary Biology 1289: 48–68. DOI: 10.1111/nyas.12082 pdf

Anderson C.N and Grether, G.F. 2013. Characterization of novel microsatellite loci for Hetaerina americana damselflies, and cross-amplification in other species. Conservation Genetics Resources 5:149-151. DOI: 10.1007/s12686-012-9755-x pdf

Chao, A., Simon-Freeman, R. and Grether, G.F. 2013. Patterns of niche partitioning and alternative reproductive strategies in an East African dung beetle assemblage. Journal of Insect Behavior 26: 525-539 DOI: 10.1007/s10905-012-9364-2 pdf

Teng, B., Dao, S., Donaldson, Z. R., and Grether, G. F. 2012. New communal roosting tradition established through experimental translocation in a Neotropical harvestman. Animal Behaviour  84:1183–1190. pdf

Kemp, D.J., Herberstein, M.E. and Grether, G.F. 2012. Unravelling the true complexity of costly colour signalling. Behavioral Ecology 23:233-236. DOI:10.1093/beheco/arr153 pdf

Anderson, C.N., Grether, G.F., Cordoba-Aguilar, A. 2012. Characterization of 12 microsatellite loci in the waterfall damselfly (Paraphlebia zoe) for use in population genetic applications. Conservation Genetics Resources 4:175–177 pdf

Deere, K.A., Grether, G.F., Sun, A. and Sinsheimer, J.S. 2012. Female mate preference explains countergradient variation in the sexual coloration of guppies (Poecilia reticulata). Proceedings of the Royal Society B. 279: 1684–1690 DOI: 10.1098/rspb.2011.2132 pdf

Anderson, C.N., Cordoba-Aguilar, A., Drury, J.P., Grether, G.F. 2011. An assessment of marking techniques for odonates in the family Calopterygidae. Entomologia Experimentalis et Applicata 141:258-261 DOI: 10.1111/j.1570-7458.2011.01185.x pdf

Grether, G.F. 2011. The neuroecology of competitor recognition. Integrative and Comparative Biology 51: 807-818. DOI: 10.1093/icb/icr060 pdf

Grether, G.F. and Kolluru, G.R. 2011. Evolutionary and plastic responses to resource availability. Ch. 6, pp. 61-71, In: Ecology and Evolution of Poeciliid Fishes. J. P. Evans, A. Pilastro, and I. Schlupp (editors). University of Chicago Press, Chicago.

Anderson, C.N. and Grether, G.F. 2011. Multiple routes to reduced interspecific territorial fighting in Hetaerina damselflies.  Behavioral Ecology 22:527-534. pdf

Anderson, C.N. and Grether, G.F. 2010. Character displacement in the fighting colors of Hetaerina damselflies.  Proceedings of the Royal Society B. 277: 3669-3675. pdf

Anderson, C.N. and Grether, G.F. 2010. Interspecific aggression and character displacement of competitor recognition in Hetaerina damselflies. Proceedings of the Royal Society B 277: 549-555. pdf.

Grether, G.F. 2010 The evolution of mate preferences, sensory biases and indicator traits. Advances in the Study of Behavior. 41: 35-76.pdf

Grether, G.F. 2010. Sexual selection and speciation. In: Encyclopedia of Animal Behavior, M. Breed and J. Moore (eds.). vol. 3, pp. 177-183, Oxford: Academic Press.

Grether, G.F. 2010. Evolution. In: The Encyclopedia of Applied Animal Behaviour & Welfare D. Mills (editor in chief), CABI Publishing, Cambridge, MA.

Grether, G.F. 2010. Selection. In: The Encyclopedia of Applied Animal Behaviour & Welfare D. Mills (editor in chief), CABI Publishing, Cambridge, MA.

Van Valkenburgh, B., Maddox, T., Funston, P.J., Mills, M.G.L., Grether, G.F., Carbone, C. 2009. Sociality in Rancho La Brea Smilodon: arguments favor “evidence” over “coincidence”. Biology Letters 5: 563-564. pdf

Kemp, D. J., Reznick, D. N., Grether, G. F., and Endler, J. A. 2009. Predicting the direction of ornament evolution in Trinidadian guppies (Poecilia reticulata). Proceedings of the Royal Society B 276: 4335-4343. pdf

Grether, G.F., Losin, N., Anderson, C.N., Okamoto, K. 2009. The role of interspecific interference competition in character displacement and the evolution of competitor recognition. Biological Reviews 84: 617–635. pdf

Kolluru, G.R., Grether, G.F., Dunlop, E., South, S. 2009. Food availability and parasite infection influence mating tactics in guppies (Poecilia reticulata). Behavioral Ecology 20: 131-173. pdf

Carbone, C., Maddox, T., Funston, P.J., Mills, M.G.L., Grether, G.F., Van Valkenburgh, B. 2008. Parallels between playbacks and Pleistocene tar seeps suggest sociality in an extinct sabretooth cat, Smilodon. Biology Letters 5: 81-85. pdf

Tatarenkov, A. Healey, C.I.M., Grether, G.F., Avise, J.C. 2008 Pronounced reproductive skew in a natural population of green swordtails, Xiphophorus helleri. Molecular Ecology 17: 4522–4534. pdf

Kemp, D. Reznick, D.N., Grether, G.F. 2008. Ornament evolution in Trinidadian guppies (Poecilia reticulata): insights from sensory processing-based analyses of entire color patterns. Biological Journal of the Linnean Society. 95: 734-747. pdf

Smith, T.B. and Grether, G.F.  2008. The importance of conserving evolutionary processes.  Pp. 85-98. In: Conservation Biology: Evolution in Action.  Scott P. Carroll and Charles W. Fox (editors).  Oxford University Press, Oxford.

Grether, G.F., Kolluru, G.R., Lin, K., Quiroz, M.A., Robertson, G., Snyder, A.J.  2008. Maternal effects of carotenoid consumption in guppies (Poecilia reticulata). Functional Ecology 22: 294-302. pdf

Smith, T.B. Milá, B, Grether, G.F., Slabbekoorn, H.,Sepil, I., Buermann, W., Saatchi, S., Pollinger, J.P. 2008 Evolutionary consequences of human disturbance in a rainforest bird species from Central Africa. Molecular Ecology 17: 58-71. pdf

Brown, D.M., Brenneman, R.A., Koepfli, K.-P., Pollinger, J.P., Milá, B. Georgiadis, N.J.. Louis Jr,, E.E. Grether, G.F., Jacobs, D.K., Wayne, R.K. 2007 Extensive population genetic structure in the giraffe.  BMC Biology 5:57. Cited by Faculty of 1000  pdf

Rainey, M.M. and Grether, G.F. 2007 Competitive mimicry: synthesis of a neglected class of mimetic relationships. Ecology 88: 2440-2448. pdf

Grether, G.F. and Donaldson, Z.R. 2007 Communal roost site selection in a Neotropical harvestman: habitat limitation versus tradition. Ethology 113: 290-300. pdf

Donaldson, Z.R. and Grether, G.F. 2007. Tradition without social learning: scent-mark-based communal roost formation in a Neotropical harvestman (Prionostemma sp.). Behavioral Ecology and Sociobiology 61: 801-809. pdf

Kolluru, G.R., Grether, G.F., Contreras, H. 2007. Environmental and genetic influences on mating strategies along a replicated food availability gradient in guppies (Poecilia reticulata). Behavioral Ecology and Sociobiology 61: 689-701. .Cited by Faculty of 1000  pdf

Kolluru, G.R., Grether, G.F., South, S., Dunlop, E., Cardinali, A., Liu, L., Carapiet, A. 2006. The effects of carotenoid and food intake on resistance to a naturally occurring parasite (Gyrodactylus turnbulli) in guppies (Poecilia reticulata). Biological Journal of the Linnean Society 89: 301-309. pdf

Kolluru, G. R., N. C. Ruiz, N. Del Cid, E. Dunlop, G. F. Grether 2006. The effects of carotenoid and food intake on caudal fin regeneration in male guppies. Journal of Fish Biology 68:1002-1012. pdf

Grether, G. F. 2005 Environmental change, phenotypic plasticity and genetic compensation. American Naturalist 166: E115-E123pdf

Grether, G. F., G. R. Kolluru, F. H. Rodd, J. de la Cerda, K. Shimazaki 2005. Carotenoid availability affects the development of a colour-based mate preference and the sensory bias to which it is genetically linked. Proceedings of the Royal Society B 272: 2181-2188. pdf

Grether, G. F., Cummings, M.E., Hudon, J. 2005. Countergradient variation in the sexual coloration of guppies (Poecilia reticulata): drosopterin synthesis balances carotenoid availability. Evolution 59: 175-188. Cited by Faculty of 1000  pdf

Kolluru, G.R. and Grether, G.F. 2004 The effects of resource availability on alternative mating tactics in guppies (Poecilia reticulata). Behavioral Ecology 16:294-300. pdf

Grether, G. F., Kolluru, G. R., Nerssisian, K. 2004. Individual color patches as multicomponent signals. Biological Reviews of the Cambridge Philosophical Society 79: 583-610. pdf

Grether, G.F., Kasahara, S., Kolluru, G.R., Cooper, E.L. 2004. Sex-specific effects of carotenoid intake on the immunological response to allografts in guppies (Poecilia reticulata). Proceedings of the Royal Society B 271: 45-49. pdf

Hudon, J., Grether, G. F., Millie, D. F.  2003. Marginal carotenoid differentiation between the sexual and general pigmentation of guppies (Poecilia reticulata) and a possible visual explanation. Physiological and Biochemical Zoology 76: 776-790. pdf

Grether, G. F., Hughes, K. A., Rodd, F. H.  2002.  Attraction to Orange: Sexiness, Not Gluttony: Response. Science 296: 847-848. pdf

Rodd, F. H., Hughes, K. A., Grether, G. F., Baril, C. T.   2002. A possible non-sexual origin of a mate preference: are male guppies mimicking fruit? Proceedings of the Royal Society B 269: 475-481. pdf

Grether, G. F., Hudon, J., Endler, J. A.  2001. Carotenoid scarcity, synthetic pteridine pigments and the evolution of sexual coloration in guppies (Poecilia reticulata). Proceedings of the Royal Society B 268:1245-1253. pdf

Grether, G.F., Millie, D.F., Bryant, M.J., Reznick, D.N., Mayea, W.  2001. Rainforest canopy cover, resource availability, and life history evolution in guppies. Ecology 82:1546-1559. pdf

Grether, G.F. 2000. Carotenoid limitation and mate preference evolution: a test of the indicator hypothesis in guppies Poecilia reticulata. Evolution 54:1712-1724. pdf

Grether, G.F. and Switzer, P.V. 2000. Mechanisms for the formation and maintenance of traditional night roost aggregations in a territorial damselfly.  Animal Behaviour 60:569-579. pdf

Switzer, P.V. and Grether, G.F. 2000. Characteristics and possible functions of traditional night roosting aggregations in rubyspot damselflies.  Behaviour 137:401-416. pdf

Grether, G. F., Hudon, J., Millie, D. F. 1999. Carotenoid limitation of sexual coloration along an environmental gradient in guppies. Proceedings of the Royal Society B 266:1317-1322. pdf

Grether, G.F. 1997. Survival cost of an intra-sexually selected ornament in a damselfly. Proceedings of the Royal Society B 264:207-210.  pdf

Grether, G.F. 1996. Sexual selection and survival selection on wing coloration and body size in the rubyspot damselfly Hetaerina americana. Evolution 50:1939-1948. pdf

Grether, G.F. 1996. Intrasexual competition alone favors a sexually dimorphic ornament in the rubyspot damselfly Hetaerina americana. Evolution  50:1949-1957. pdf

Grether, G.F. & Grey, R.M. 1996. Novel cost of a sexually selected trait in the rubyspot damselfly Hetaerina americana: conspicuousness to prey. Behavioral Ecology 7:465-473. pdf

Grether, G.F. 1995. Genetic correlation complicates the interpretation of phylogenetic transitions in sexually selected traits. Behavioral Ecology 6:349. pdf

Grether, G.F., Palombit, R.A., Rodman, P.S. 1992. International Journal of Primatology 13:1–17. pdf

Hamilton, W.J. III, Grether, G.F., Easley, S. 1992. Test of a recapture radio collar suitable for non-human primates. Contemporary Topics in Laboratory Animal Science 31:8–9.

Mitani, J.C., Grether, G.F., Rodman, P.S., Priatna, D. 1991. Associations among wild orangutans: sociality, passive aggregation or chance? Animal Behaviour 42:33–46. pdf

Grether, G.F. and Weaver, A.M. 1990. What are sisters good for? Nature 345:392. pdf